俞立波 , 鲁统宇 , 孙建明 . 预期视角下我国大豆价格影响因素研究 [J ] . 中国物价 , 2024 ( 10 ): 13 - 18 .

YU L B , LU T Y , SUN J M . Study on the influencing factors of domestic soybean price from the perspective of expectancy [J ] . China Price Journal , 2024 ( 10 ): 13 - 18 .

ABDELGHANY A M , ZHANG S R , AZAM M , et al . Profiling of seed fatty acid composition in 1025 Chinese soybean accessions from diverse ecoregions [J ] . The Crop Journal , 2020 , 8 ( 4 ): 635 - 644 .

WU Y , ZHOU R S , WANG Z G , et al . The effect of refining process on the physicochemical properties and micronutrients of rapeseed oils [J ] . PLoS One , 2019 , 14 ( 3 ): e0212879 .

LI M H , LUO J N , NAWAZ M A , et al . Phytochemistry, bioaccessibility, and bioactivities of sesame seeds: an overview [J ] . Food Reviews International , 2024 , 40 ( 1 ): 309 - 335 .

MUHAMMAD ANJUM F , NADEEM M , ISSA KHAN M , et al . Nutritional and therapeutic potential of sunflower seeds: a review [J ] . British Food Journal , 2012 , 114 ( 4 ): 544 - 552 .

CAO H P , GONG W F , RONG J , et al . Editorial: Woody oil crops: key trait formation and regulation [J ] . Frontiers in Plant Science , 2023 , 14 : 1328990 .

ZHANG F H , LI Z , ZHOU J Q , et al . Comparative study on fruit development and oil synthesis in two cultivars of Camellia oleifera [J ] . BMC Plant Biology , 2021 , 21 ( 1 ): 348 .

YI M Y , YOU Y , ZHANG Y R , et al . Highly valuable fish oil: formation process, enrichment, subsequent utilization, and storage of eicosapentaenoic acid ethyl esters [J ] . Molecules , 2023 , 28 ( 2 ): 672 .

MISHRA B , AKHILA MV A , THOMAS A , et al . Formulated therapeutic products of animal fats and oils: future prospects of zootherapy [J ] . International Journal of Pharmaceutical Investigation , 2020 , 10 ( 2 ): 112 - 116 .

ATHENAKI M , GARDELI C , DIAMANTOPOULOU P , et al . Lipids from yeasts and fungi: physiology, production and analytical considerations [J ] . Journal of Applied Microbiology , 2018 , 124 ( 2 ): 336 - 367 .

ALVAREZ H M , HERNÁNDEZ M A , LANFRANCONI M P , et al . Rhodococcus as biofactories for microbial oil production [J ] . Molecules , 2021 , 26 ( 16 ): 4871 .

BLAZECK J , HILL A , LIU L Q , et al . Harnessing Yarrowia lipolytica lipogenesis to create a platform for lipid and biofuel production [J ] . Nature Communications , 2014 , 5 : 3131 .

徐鹏 . 纪念王义翘教授: 解脂耶氏酵母替代植物油脂的技术瓶颈及展望 [J ] . 合成生物学 , 2021 , 2 ( 4 ): 509 - 527 .

XU P . In memory of Prof. Daniel I. C. Wang: engineering Yarrowia lipolytica for the production of plant-based lipids: technical constraints and perspectives for a sustainable cellular agriculture economy [J ] . Synthetic Biology Journal , 2021 , 2 ( 4 ): 509 - 527 .

LARROUDE M , ROSSIGNOL T , NICAUD J M , et al . Synthetic biology tools for engineering Yarrowia lipolytica [J ] . Biotechnology Advances , 2018 , 36 ( 8 ): 2150 - 2164 .

CHRISTEN S , SAUER U . Intracellular characterization of aerobic glucose metabolism in seven yeast species by 13 C flux analysis and metabolomics [J ] . FEMS Yeast Research , 2011 , 11 ( 3 ): 263 - 272 .

SPAGNUOLO M , YAGUCHI A , BLENNER M . Oleaginous yeast for biofuel and oleochemical production [J ] . Current Opinion in Biotechnology , 2019 , 57 : 73 - 81 .

XUE S J , CHI Z , ZHANG Y , et al . Fatty acids from oleaginous yeasts and yeast-like fungi and their potential applications [J ] . Critical Reviews in Biotechnology , 2018 , 38 ( 7 ): 1049 - 1060 .

YAEGASHI J , KIRBY J , ITO M , et al . Rhodosporidium toruloides : a new platform organism for conversion of lignocellulose into terpene biofuels and bioproducts [J ] . Biotechnology for Biofuels , 2017 , 10 : 241 .

LI Y H , ZHAO Z B , BAI F W . High-density cultivation of oleaginous yeast Rhodosporidium toruloides Y4 in fed-batch culture [J ] . Enzyme and Microbial Technology , 2007 , 41 ( 3 ): 312 - 317 .

GÖRNER C , REDAI V , BRACHARZ F , et al . Genetic engineering and production of modified fatty acids by the non-conventional oleaginous yeast Trichosporon oleaginosus ATCC 20509 [J ] . Green Chemistry , 2016 , 18 ( 7 ): 2037 - 2046 .

YAGUCHI A , ROBINSON A , MIHEALSICK E , et al . Metabolism of aromatics by Trichosporon oleaginosus while remaining oleaginous [J ] . Microbial Cell Factories , 2017 , 16 ( 1 ): 206 .

TOTANI N , WATANABE A , OBA K . An improved method of arachidonic acid production by Mortierella alpina [J ] . Journal of Japan Oil Chemists’ Society , 1987 , 36 ( 5 ): 328 - 331 .

TAKENO S , SAKURADANI E , TOMI A , et al . Transformation of oil-producing fungus, Mortierella alpina 1S-4, using Zeocin, and application to arachidonic acid production [J ] . Journal of Bioscience and Bioengineering , 2005 , 100 ( 6 ): 617 - 622 .

SAKURADANI E , ANDO A , SHIMIZU S , et al . Metabolic engineering for the produc tion of polyunsaturated fatty acids by oleaginous fungus Mortierella alpina 1S-4 [J ] . Journal of Bioscience and Bioengineering , 2013 , 116 ( 4 ): 417 - 422 .

ZHANG X Y , LI B , HUANG B C , et al . Production, biosynthesis, and commercial applications of fatty acids from oleaginous fungi [J ] . Frontiers in Nutrition , 2022 , 9 : 873657 .

AHMAD FAZILI A B , SHAH A M , ZAN X Y , et al . Mucor circinelloides : a model organism for oleaginous fungi and its potential applications in bioactive lipid production [J ] . Microbial Cell Factories , 2022 , 21 ( 1 ): 29 .

HAN X , LI Z H , WEN Y , et al . Overproduction of docosahexaenoic acid in Schizochytrium sp. through genetic engineering of oxidative stress defense pathways [J ] . Biotechnology for Biofuels , 2021 , 14 ( 1 ): 70 .

BARCLAY W , WEAVER C , METZ J , et al . Development of a docosahexaenoic acid production technology using Schizochytrium : historical perspective and update [M/OL ] // COHEN Z, RATLEDGE C. 2nd Edition . Single cell oils. Amsterdam: Elsevier , 2010 : 75 - 96 . ( 2015-09-18)[2025-03-01] . https://doi.org/10.1016/B978-1-893997-73-8.50008-6 https://doi.org/10.1016/B978-1-893997-73-8.50008-6 .

MATOS Â P , FELLER R , MOECKE E H S , et al . Chemical characterization of six microalgae with potential utility for food application [J ] . Journal of the American Oil Chemists’ Society , 2016 , 93 ( 7 ): 963 - 972 .

RIZWAN M , MUJTABA G , MEMON S A , et al . Exploring the potential of microalgae for new biotechnology applications and beyond: a review [J ] . Renewable and Sustainable Energy Reviews , 2018 , 92 : 394 - 404 .

XUE Z H , YU Y , YU W C , et al . Development prospect and preparation technology of edible oil from microalgae [J ] . Frontiers in Marine Science , 2020 , 7 : 402 .

KIM H M , CHAE T U , CHOI S Y , et al . Engineering of an oleaginous bacterium for the production of fatty acids and fuels [J ] . Nature Chemical Biology , 2019 , 15 ( 7 ): 721 - 729 .

GIENTKA I , GADASZEWSKA M , BŁAŻEJAK S , et al . Evaluation of lipid biosynthesis ability by Rhodotorula and Sporobolomyces strains in medium with glycerol [J ] . European Food Research and Technology , 2017 , 243 ( 2 ): 275 - 286 .

GARG S , WU H , CLOMBURG J M , et al . Bioconversion of methane to C-4 carboxylic acids using carbon flux through acetyl-CoA in engineered Methylomicrobium buryatense 5GB1C [J ] . Metabolic Engineering , 2018 , 48 : 175 - 183 .

KIKUKAWA H , SAKURADANI E , ANDO A , et al . Arachidonic acid production by the oleaginous fungus Mortierella alpina 1S-4: a review [J ] . Journal of Advanced Research , 2018 , 11 : 15 - 22 .

JIN H , ZHANG H , ZHOU Z W , et al . Ultrahigh-cell-density heterotrophic cultivation of the unicellular green microalga Scenedesmus acuminatus and application of the cells to photoautotrophic culture enhance biomass and lipid production [J ] . Biotechnology and Bioengineering , 2020 , 117 ( 1 ): 96 - 108 .

JOVANOVIC S , DIETRICH D , BECKER J , et al . Microbial production of polyunsaturated fatty acids: high-value ingredients for aquafeed, superfoods, and pharmaceuticals [J ] . Current Opinion in Biotechnology , 2021 , 69 : 199 - 211 .

SU H , SHI P H , SHEN Z S , et al . High-level production of nervonic acid in the oleaginous yeast Yarrowia lipolytica by systematic metabolic engineering [J ] . Communications Biology , 2023 , 6 : 1125 .

PFLEGER B F , GOSSING M , NIELSEN J . Metabolic engineering strategies for microbial synthesis of oleochemicals [J ] . Metabolic Engineering , 2015 , 29 : 1 - 11 .

YAN Q , PFLEGER B F . Revisiting metabolic engineering strategies for microbial synthesis of oleochemicals [J ] . Metabolic Engineering , 2020 , 58 : 35 - 46 .

KHAN I , HUSSAIN M , JIANG B Z , et al . Omega-3 long-chain polyunsaturated fatty acids: metabolism and health implications [J ] . Progress in Lipid Research , 2023 , 92 : 101255 .

LENNEN R M , PFLEGER B F . Microbial production of fatty acid-derived fuels and chemicals [J ] . Current Opinion in Biotechnology , 2013 , 24 ( 6 ): 1044 - 1053 .

RAHIM M A , AYUB H , SEHRISH A , et al . Essential components from plant source oils: a review on extraction, detection, identification, and quantification [J ] . Molecules , 2023 , 28 ( 19 ): 6881 .

GALLEGO-GARCÍA M , SUSMOZAS A , NEGRO M J , et al . Challenges and prospects of yeast-based microbial oil production within a biorefinery concept [J ] . Microbial Cell Factories , 2023 , 22 ( 1 ): 246 .

UKEGBU P O , ONWUZURUIKE U A , OBASI N E . Production of edible oil from microorganisms [M ] //BABALOLA O O. Food security and safety . Cham : Springer International Publishing , 2021 : 563 - 592 . ( 2021-09-21)[2025-03-01] . https://doi.org/10.1007/978-3-030-50672-8_29 https://doi.org/10.1007/978-3-030-50672-8_29 .

JIN H , YANG X , ZHAO H B , et al . Genetic analysis of protein content and oil content in soybean by genome-wide association study [J ] . Frontiers in Plant Science , 2023 , 14 : 1182771 .

DUAN Z B , LI Q , WANG H , et al . Genetic regulatory networks of soybean seed size, oil and protein contents [J ] . Frontiers in Plant Science , 2023 , 14 : 1160418 .

QIAO K J , IMAM ABIDI S H , LIU H J , et al . Engineering lipid overproduction in the oleaginous yeast Yarrowia lipolytica [J ] . Metabolic Engineering , 2015 , 29 : 56 - 65 .

QIAO K J , WASYLENKO T M , ZHOU K , et al . Lipid production in Yarrowia lipolytica is maximized by engineering cytosolic redox metabolism [J ] . Nature Biotechnology , 2017 , 35 ( 2 ): 173 - 177 .

CORTÉS-PEÑA Y R , WOODRUFF W , BANERJEE S , et al . Integration of plant and microbial oil processing at oilcane biorefineries for more sustainable biofuel production [J ] . GCB Bioenergy , 2024 , 16 ( 11 ): e13183 .

LUDWICZAK A , ZIELIŃSKI T , SIBIŃSKA E , et al . Comparative analysis of microbial contamination in diesel fuels using MALDI-TOF MS [J ] . Scientific Reports , 2025 , 15 : 4525 .

GHULAM KADIR A P . Oil palm economic performance in Malaysia and R&D progress in 2019 [J ] . Journal of Oil Palm Research , 2020 : 159 - 190 .

PARK S B , LEE Y R , YUN J H , et al . Towards maximizing biomass and lipid productivity: high-throughput screening assay for prospecting heterotrophic growth for new microalgal isolates [J ] . Microbial Cell Factories , 2024 , 23 ( 1 ): 299 .

MEIJAARD E M , VIRAH-SAWMY M , NEWING H S , et al . Exploring the future of vegetable oils [M ] . Gland, Switzerland : IUCN; SNSB , 2024 .

SONG X P , HANSEN M C , POTAPOV P , et al . Massive soybean expansion in South America since 2000 and implications for conservation [J ] . Nature Sustainability , 2021 , 4 ( 9 ): 784 - 792 .

CLEMENTE T E , CAHOON E B . Soybean oil: genetic approaches for modification of functionality and total content [J ] . Plant Physiology , 2009 , 151 ( 3 ): 1030 - 1040 .

LARDIZABAL K , EFFERTZ R , LEVERING C , et al . Expression of Umbelopsis ramanniana DGAT2A in seed increases oil in soybean [J ] . Plant Physiology , 2008 , 148 ( 1 ): 89 - 96 .

SONG Q X , LI Q T , LIU Y F , et al . Soybean GmbZIP123 gene enhances lipid content in the seeds of transgenic Arabidopsis plants [J ] . Journal of Experimental Botany , 2013 , 64 ( 14 ): 4329 - 4341 .

WANG H W , ZHANG B , HAO Y J , et al . The soybean Dof-type transcription factor genes, GmDof4 and GmDof11, enhance lipid content in the seeds of transgenic Arabidopsis plants [J ] . The Plant Journal , 2007 , 52 ( 4 ): 716 - 729 .

LIU Y F , LI Q T , LU X , et al . Soybean GmMYB73 promotes lipid accumulation in transgenic plants [J ] . BMC Plant Biology , 2014 , 14 ( 1 ): 73 .

刘虹洁 , 王金星 , 刘昭军 , 等 . 大豆种子蛋白和油脂含量调控的研究进展 [J ] . 热带亚热带植物学报 , 2022 , 30 ( 6 ): 791 - 800 .

LIU H J , WANG J X , LIU Z J , et al . Research progress on protein and oil contents of soybean seeds [J ] . Journal of Tropical and Subtropical Botany , 2022 , 30 ( 6 ): 791 - 800 .

TOKEL D , ERKENCIOGLU B N . Production and trade of oil crops, and their contribution to the world economy [M/OL ] // TOMBULOGLU H, UNVER T, TOMBULOGLU G, et al. Oil crop genomics . Cham : Springer International Publishing , 2021 : 415 - 427 . ( 2021-09-21)[2025-03-01] . https://doi.org/10.1007/978-3-030-70420-9_20 https://doi.org/10.1007/978-3-030-70420-9_20 .

CHEW T L , BHATIA S . Catalytic processes towards the production of biofuels in a palm oil and oil palm biomass-based biorefinery [J ] . Bioresource Technology , 2008 , 99 ( 17 ): 7911 - 7922 .

TAN K T , LEE K T , MOHAMED A R , et al . Palm oil: addressing issues and towards sustainable development [J ] . Renewable and Sustainable Energy Reviews , 2009 , 13 ( 2 ): 420 - 427 .

BELLOU S , TRIANTAPHYLLIDOU I E , AGGELI D , et al . Microbial oils as food additives: recent approaches for improving microbial oil production and its polyunsaturated fatty acid content [J ] . Current Opinion in Biotechnology , 2016 , 37 : 24 - 35 .

SCHÖRKEN U , KEMPERS P . Lipid biotechnology: industrially relevant production processes [J ] . European Journal of Lipid Science and Technology , 2009 , 111 ( 7 ): 627 - 645 .

RATLEDGE C . Yeasts, moulds, algae and bacteria as sources of lipids [M/OL ] //KAMEL B S, KAKUDA Y. Technological advances in improved and alternative sources of lipids . Boston, MA : Springer , 1994 : 235 - 291 [2025-03-01] . https://doi.org/10.1007/978-1-4615-2109-9_9 https://doi.org/10.1007/978-1-4615-2109-9_9 .

HASSANE A M A , ELDIEHY K S H , SAHA D , et al . Oleaginous fungi: a promising source of biofuels and nutraceuticals with enhanced lipid production strategies [J ] . Archives of Microbiology , 2024 , 206 ( 7 ): 338 .

MENG X , YANG J M , XU X , et al . Biodiesel production from oleaginous microorganisms [J ] . Renewable Energy , 2009 , 34 ( 1 ): 1 - 5 .

赵宗保 , 胡翠敏 . 能源微生物油脂技术进展 [J ] . 生物工程学报 , 2011 , 27 ( 3 ): 427 - 435 .

ZHAO Z B , HU C M . Progress in bioenergy-oriented microbial lipid technology [J ] . Chinese Journal of Biotechnology , 2011 , 27 ( 3 ): 427 - 435 .

SAENGE C , CHEIRSILP B , SUKSAROGE T T , et al . Potential use of oleaginous red yeast Rhodotorula glutinis for the bioconversion of crude glycerol from biodiesel plant to lipids and carotenoids [J ] . Process Biochemistry , 2011 , 46 ( 1 ): 210 - 218 .

LIANG Y N , TANG T Y , SIDDARAMU T , et al . Lipid production from sweet sorghum bagasse through yeast fermentation [J ] . Renewable Energy , 2012 , 40 ( 1 ): 130 - 136 .

BANERJEE S , SINGH V . Economic and environmental bottlenecks in the industrial-scale production of lipid-derived biofuels from oleaginous yeasts: a review of the current trends and future prospects [J ] . GCB Bioenergy , 2024 , 16 ( 7 ): e13173 .

QADEER S , KHALID A , MAHMOOD S , et al . Utilizing oleaginous bacteria and fungi for cleaner energy production [J ] . Journal of Cleaner Production , 2017 , 168 : 917 - 928 .

INFANTE E G , GOMIDE F T F , SECCHI A R , et al . Diesel production from lignocellulosic residues: trends, challenges and opportunities [J ] . Biofuels, Bioproducts and Biorefining , 2024 , 18 ( 5 ): 1711 - 1738 .

ZHANG B , KHUSHIK F A , ZHAN B R , et al . Transformation of lignocellulose to starch-like carbohydrates by organic acid-catalyzed pretreatment and biological detoxification [J ] . Biotechnology and Bioengineering , 2021 , 118 ( 10 ): 4105 - 4118 .

LIU G , ZHANG Q , LI H X , et al . Dry biorefining maximizes the potentials of simultaneous saccharification and co-fermentation for cellulosic ethanol production [J ] . Biotechnology and Bioengineering , 2018 , 115 ( 1 ): 60 - 69 .

LIU Q , LU M P , JIN C , et al . Ultra-centrifugation force in adaptive evolution changes the cell structure of oleaginous yeast Trichosporon cutaneum into a favorable space for lipid accumulation [J ] . Biotechnology and Bioengineering , 2022 , 119 ( 6 ): 1509 - 1521 .

COTTON C A , EDLICH-MUTH C , BAR-EVEN A . Reinforcing carbon fixation: CO 2 reduction replacing and supporting carboxylation [J ] . Current Opinion in Biotechnology , 2018 , 49 : 49 - 56 .

LI A P , CAO X P , FU R Z , et al . Biocatalysis of CO 2 and CH 4 : key enzymes and challenges [J ] . Biotechnology Advances , 2024 , 72 : 108347 .

赵亮 , 李振帅 , 付丽平 , 等 . 生物转化一碳化合物原料产油脂与单细胞蛋白研究进展 [J ] . 合成生物学 , 2024 , 5 ( 6 ): 1300 - 1318 .

ZHAO L , LI Z S , FU L P , et al . Progress in biomanufacturing of lipids and single cell protein from one-carbon compounds [J ] . Synthetic Biology Journal , 2024 , 5 ( 6 ): 1300 - 1318 .

KINNEY A J , CAHOON E B , DAMUDE H G , et al . Production of very long chain polyunsaturated fatty acids in oilseed plants : US 13/044984 [P ] . 2011-11-03 .

RUIZ-LÓPEZ N , SAYANOVA O , NAPIER J A , et al . Metabolic engineering of the omega-3 long chain polyunsaturated fatty acid biosynthetic pathway into transgenic plants [J ] . Journal of Experimental Botany , 2012 , 63 ( 7 ): 2397 - 2410 .

HUDSON K A , HUDSON M E . Genetic variation for seed oil biosynthesis in soybean [J ] . Plant Molecular Biology Reporter , 2021 , 39 ( 4 ): 700 - 709 .

ZHOU B , FEI W J , YANG S Q , et al . Alteration of the fatty acid composition of Brassica napus L. via overexpression of phospholipid: diacylglycerol acyltransferase 1 from Sapium sebiferum (L. ) Roxb [J ] . Plant Science , 2020 , 298 : 110562 .

FENYK S , WOODFIELD H K , ROMSDAHL T B , et al . Overexpression of phospholipid: diacylglycerol acyltransferase in Brassica napus results in changes in lipid metabolism and oil accumulation [J ] . The Biochemical Journal , 2022 , 479 ( 6 ): 805 - 823 .

WU D , ZHANG K , LI C Y , et al . Genome-wide comprehensive characterization and transcriptomic analysis of AP2 / ERF gene family revealed its role in seed oil and ALA formation in Perilla ( Perilla frutescens ) [J ] . Gene , 2023 , 889 : 147808 .

SINCLAIR A J , JAYASOORIYA A . Nutritional aspects of single cell oils: applications of arachidonic acid and docosahexaenoic acid oils [M/OL ] //COHEN Z, RATLEDGE C. 2nd Edition . Single cell oils. Amsterdam: Elsevier , 2010 : 351 - 368 . ( 2015-09-18)[2025-03-01] . https://doi.org/10.1016/B978-1-893997-73-8.50020-7 https://doi.org/10.1016/B978-1-893997-73-8.50020-7 .

XIE D M , JACKSON E N , ZHU Q . Sustainable source of omega-3 eicosapentaenoic acid from metabolically engineered Yarrowia lipolytica : from fundamental research to commercial production [J ] . Applied Microbiology and Biotechnology , 2015 , 99 ( 4 ): 1599 - 1610 .

XUE Z X , SHARPE P L , HONG S P , et al . Production of omega-3 eicosapentaenoic acid by metabolic engineering of Yarrowia lipolytica [J ] . Nature Biotechnology , 2013 , 31 ( 8 ): 734 - 740 .

DU F , LI Z J , LI X , et al . Optimizing multicopy chromosomal integration for stable high-performing strains [J ] . Nature Chemical Biology , 2024 , 20 ( 12 ): 1670 - 1679 .

CUI J , CHEN H Q , TANG X , et al . Δ6 Fatty acid desaturases in polyunsaturated fatty acid biosynthesis: insights into the evolution, function with substrate specificities and biotechnological use [J ] . Applied Microbiology and Biotechnology , 2020 , 104 ( 23 ): 9947 - 9963 .

CHAKRABORTY P , KUMAR R , KARN S , et al . Recent trends in metabolic engineering for microbial production of value-added natural products [J ] . Biochemical Engineering Journal , 2025 , 213 : 109537 .

RANI H , SHARMA S , BALA M . Technologies for extraction of oil from oilseeds and other plant sources in retrospect and prospects: a review [J ] . Journal of Food Process Engineering , 2021 , 44 ( 11 ): e13851 .

ZENG W Q , LIU X D , CHAO Y , et al . The effect of extraction methods on the components and quality of Camellia oleifera oil: focusing on the flavor and lipidomics [J ] . Food Chemistry , 2024 , 447 : 139046 .

ZHANG F , ZHU F , CHEN B L , et al . Composition, bioactive substances, extraction technologies and the influences on characteristics of Camellia oleifera oil: a review [J ] . Food Research International , 2022 , 156 : 111159 .

KAPOORE R V , BUTLER T O , PANDHAL J , et al . Microwave-assisted extraction for microalgae: from biofuels to biorefinery [J ] . Biology , 2018 , 7 ( 1 ): 18 .

GRAJZER M , SZMALCEL K , KUŹMIŃSKI Ł , et al . Characteristics and antioxidant potential of cold-pressed oils-possible strategies to improve oil stability [J ] . Foods , 2020 , 9 ( 11 ): 1630 .

SYMONIUK E , WRONIAK M , NAPIÓRKOWSKA K , et al . Oxidative stability and antioxidant activity of selected cold-pressed oils and oils mixtures [J ] . Foods , 2022 , 11 ( 11 ): 1597 .

POSTMA P R , MIRON T L , OLIVIERI G , et al . Mild disintegration of the green microalgae Chlorella vulgaris using bead milling [J ] . Bioresource Technology , 2015 , 184 : 297 - 304 .

MEULLEMIESTRE A , BREIL C , ABERT-VIAN M , et al . Microwave, ultrasound, thermal treatments, and bead milling as intensification techniques for extraction of lipids from oleaginous Yarrowia lipolytica yeast for a biojetfuel application [J ] . Bioresource Technology , 2016 , 211 : 190 - 199 .

EKPENI L E N , BENYOUNIS K Y , NKEM-EKPENI F F , et al . Underlying factors to consider in improving energy yield from biomass source through yeast use on high-pressure homogenizer (hph) [J ] . Energy , 2015 , 81 : 74 - 83 .

LIU C Z , WUFUER A , KONG L P , et al . Organic solvent extraction-assisted catalytic hydrothermal liquefaction of algae to bio-oil [J ] . RSC Advances , 2018 , 8 ( 55 ): 31717 - 31724 .

WRONA O , RAFIŃSKA K , MOŻEŃSKI C , et al . Supercritical fluid extraction of bioactive compounds from plant materials [J ] . Journal of AOAC International , 2017 , 100 ( 6 ): 1624 - 1635 .

DANLAMI J M , ARSAD A , AHMAD ZAINI M A , et al . A comparative study of various oil extraction techniques from plants [J ] . Reviews in Chemical Engineering , 2014 , 30 ( 6 ): 605 - 626 .

WANG C X , DUAN Z H , FAN L P , et al . Supercritical CO 2 fluid extraction of Elaeagnus mollis Diels seed oil and its antioxidant ability [J ] . Molecules , 2019 , 24 ( 5 ): 911 .

KHOO H E , AZLAN A , ABD KADIR N A A . Fatty acid profile, phytochemicals, and other substances in Canarium odontophyllum fat extracted using supercritical carbon dioxide [J ] . Frontiers in Chemistry , 2019 , 7 : 5 .

CHENG C H , DU T B , PI H C , et al . Comparative study of lipid extraction from microalgae by organic solvent and supercritical CO 2 [J ] . Bioresource Technology , 2011 , 102 ( 21 ): 10151 - 10153 .

GAO Y H , DING Z S , LIU Y F , et al . Aqueous enzymatic extraction: a green, environmentally friendly and sustainable oil extraction technology [J ] . Trends in Food Science & Technology , 2024 , 144 : 104315 .

JIN G J , YANG F , HU C M , et al . Enzyme-assisted extraction of lipids directly from the cu lture of the oleaginous yeast Rhodosporidium toruloides [J ] . Bioresource Technology , 2012 , 111 : 378 - 382 .

张珺璐 , 吕力婷 , 梁世玉 , 等 . 基于酵母油脂技术的生物质炼制 [J ] . 中国科学:化学 , 2025 , 55 ( 1 ): 28 - 36 .

ZHANG J L , LYU L T , LIANG S Y , et al . Yeast lipid technology for biomass refinery [J ] . Scientia Sinica Chimica , 2025 , 55 ( 1 ): 28 - 36 .

CRAVOTTO G , BOFFA L , MANTEGNA S , et al . Improved extraction of vegetable oils under high-intensity ultrasound and/or microwaves [J ] . Ultrasonics Sonochemistry , 2008 , 15 ( 5 ): 898 - 902 .

THILAKARATHNA R C N , SIOW L F , TANG T K , et al . A review on application of ultrasound and ultrasound assisted technology for seed oil extraction [J ] . Journal of Food Science and Technology , 2023 , 60 ( 4 ): 1222 - 1236 .

MATEI P L , DELEANU I , BREZOIU A M , et al . Ultrasound-assisted extraction of blackberry seed oil: optimization and oil characterization [J ] . Molecules , 2023 , 28 ( 6 ): 2486 .

ZHANG X L , YAN S , TYAGI R D , et al . Ultrasonication assisted lipid extraction from oleaginous microorganisms [J ] . Bioresource Technology , 2014 , 158 : 253 - 261 .

LU Y C , XIONG R X , TANG Y C , et al . An overview of the detection methods to the edible oil oxidation degree: recent progress, challenges, and perspectives [J ] . Food Chemistry , 2025 , 463 : 141443 .

ECKER J , SCHERER M , SCHMITZ G , et al . A rapid GC-MS method for quantification of positional and geometric isomers of fatty acid methyl esters [J ] . Journal of Chromatography B, Analytical Technologies in the Biomedical and Life Sciences , 2012 , 897 : 98 - 104 .

PATEL M K , DAS S , THAKUR J K . GC-MS-based analysis of methanol: chloroform-extracted fatty acids from plant tissues [J ] . Bio-protocol , 2018 , 8 ( 18 ): e3014 .

ZHONG H Q , CHAI J Y , YU C L , et al . Rapid detection of oil content in Camellia oleifera kernels based on hyperspectral imaging and machine learning [J ] . Journal of Food Composition and Analysis , 2025 , 137 : 106899 .

YUAN W D , ZHOU H P , ZHANG C , et al . Prediction of oil content in Camellia oleifera seeds based on deep learning and hyperspectral imaging [J ] . Industrial Crops and Products , 2024 , 222 : 119662 .

SONG A R , WANG C Y , WEN W L , et al . Predicting the oil content of individual corn kernels combining NIR-HSI and multi-stage parameter optimization techniques [J ] . Food Chemistry , 2024 , 461 : 140932 .

HE Y H , WANG X X , MA B , et al . Ramanome technology platform for label-free screening and sorting of microbial cell factories at single-cell resolution [J ] . Biotechnology Advances , 2019 , 37 ( 6 ): 107388 .

PRADO E , EKLOUH-MOLINIER C , ENEZ F , et al . Prediction of fatty acids composition in the rainbow trout Oncorhynchus mykiss by using Raman micro-spectroscopy [J ] . Analytica Chimica Acta , 2022 , 1191 : 339212 .

SASAKI R , TODA S , SAKAMOTO T , et al . Simultaneous imaging and characterization of polyunsaturated fatty acids, carotenoids, and microcrystalline guanine in single Aurantiochytrium limacinum cells with linear and nonlinear Raman microspectroscopy [J ] . The Journal of Physical Chemistry B , 2023 , 127 ( 12 ): 2708 - 2718 .